LLPS-Art-1836
CLPC2

▼ OVERVIEW


Status: Reviewed
Protein Name: Chaperone protein ClpC2, chloroplastic; ATP-dependent Clp protease ATP-binding subunit ClpC homolog 2; AtClpC; Casein lytic proteinase C2
Gene Name: CLPC2, HSP93-III, At3g48870, T21J18.140
Ensembl Gene: AT3G48870
Ensembl Protein: AT3G48870.4
Organism: Arabidopsis thaliana
Taxa ID: 3702
LLPS Type: Others


▼ PROPERTY



——— Disorder propensity (calculated by IUPred2A)

▼ Classification


Condensates:
CondensateDescriptionTissue/CellPMIDs
Nucleolus
"...We identified 1602 proteins in the nucleolar and 2544 proteins in the nuclear fraction with an overlap of 1429 proteins."
Arabidopsis thaliana cells26980300

▼ FUNCTION


Molecular chaperone (PubMed:15304652, PubMed:21737456, PubMed:24599948). May act as a suppressor of FtsH-mediated thylakoid membrane biogenesis and may enhance photoinhibition (PubMed:15304652). Seems not involved in chloroplastic protein import (PubMed:15304652). Probable component of the TIC-associated stromal import motor involved in inner membrane translocation (PubMed:17376159). Has an ATPase activity, but no ADPase activity (PubMed:21737456). Interacts with transit peptides with a positional preference (PubMed:21737456, PubMed:22545953). Localization of the signal sequence at the N-terminal end of a protein seems mandatory for interaction to take place (PubMed:22545953).

▼ SEQUENCE


Protein Sequence (FASTA)
1     MAWSIALLTP  PFFGPGRHVQ  AKEYREPRGC  VMKMSSLKAP  VLRIQATEYR  EPRGRVKMMS  60
61    SLQAPLLTIQ  SFSGLRAPSA  LDYLGRPSPG  FLVKYKLAKS  SGREKASRCV  PKAMFERFTE  120
121   KAIKVIMLSQ  EEARRLGHNF  VGTEQILLGL  IGEGTGIAAK  VLKSMGINLK  DSRVEVEKII  180
181   GRGSGFVAVE  IPFTPRAKRV  LELSLEEARQ  LGHNYIGSEH  LLLGLLREGE  GVAARVLENL  240
241   GADPSNIRTQ  VIRMVGENNE  VTASVGGGSS  GNSKMPTLEE  YGTNLTKLAE  EGKLDPVVGR  300
301   QPQIERVVQI  LARRTKNNPC  LIGEPGVGKT  AIAEGLAQRI  ASGDVPETIE  GKTVITLDMG  360
361   LLVAGTKYRG  EFEERLKKLM  EEIRQSDEII  LFIDEVHTLI  GAGAAEGAID  AANILKPALA  420
421   RGELQCIGAT  TIDEYRKHIE  KDPALERRFQ  PVKVPEPTVE  EAIQILQGLR  ERYEIHHKLR  480
481   YTDEALVAAA  QLSHQYISDR  FLPDKAIDLI  DEAGSRVRLR  HAQLPEEARE  LEKQLRQITK  540
541   EKNEAVRSQD  FEMAGSHRDR  EIELKAEIAN  VLSRGKEVAK  AENEAEEGGP  TVTESDIQHI  600
601   VATWTGIPVE  KVSSDESSRL  LQMEQTLHTR  VIGQDEAVKA  ISRAIRRARV  GLKNPNRPIA  660
661   SFIFSGPTGV  GKSELAKALA  AYYFGSEEAM  IRLDMSEFME  RHTVSKLIGS  PPGYVGYTEG  720
721   GQLTEAVRRR  PYTLVLFDEI  EKAHPDVFNM  MLQILEDGRL  TDSKGRTVDF  KNTLLIMTSN  780
781   VGSSVIEKGG  RRIGFDLDHD  EKDSSYNRIK  SLVTEELKQY  FRPEFLNRLD  EMIVFRQLTK  840
841   LEVKEIADIM  LKEVVARLEV  KEIELQVTER  FKERVVDEGF  DPSYGARPLR  RAIMRLLEDS  900
901   MAEKMLSRDI  KEGDSVIVDV  DAEGSVVVLS  GTTGRVGGFA  AEEAMEDPIP  IL  952
Nucleotide CDS Sequence (FASTA)
1     ATGGCTTGGT  CGATAGCTCT  TCTTACACCT  CCTTTTTTTG  GTCCCGGAAG  GCATGTGCAG  60
61    GCCAAAGAAT  ACAGGGAACC  AAGAGGGTGT  GTGATGAAGA  TGAGCAGTTT  AAAAGCACCT  120
121   GTTCTGAGAA  TTCAGGCCAC  AGAATACAGA  GAACCAAGAG  GGCGTGTGAA  GATGATGTCC  180
181   AGTTTACAAG  CACCTCTTCT  GACAATTCAG  AGCTTCTCAG  GGTTAAGGGC  CCCCAGTGCA  240
241   TTAGATTATT  TGGGAAGGCC  TAGTCCAGGT  TTCCTTGTTA  AGTATAAACT  TGCAAAATCA  300
301   TCTGGGAGAG  AAAAAGCTAG  CCGATGTGTA  CCCAAAGCAA  TGTTTGAGCG  TTTTACCGAG  360
361   AAAGCAATTA  AGGTCATAAT  GCTGTCTCAA  GAGGAAGCTC  GGAGACTTGG  CCATAACTTT  420
421   GTTGGGACTG  AGCAAATACT  GTTGGGTCTA  ATTGGAGAAG  GGACTGGGAT  TGCCGCCAAG  480
481   GTTCTTAAAT  CCATGGGGAT  CAATCTTAAA  GATTCACGCG  TGGAAGTAGA  AAAGATAATT  540
541   GGGAGAGGCA  GTGGATTCGT  GGCAGTGGAG  ATTCCATTTA  CTCCTCGCGC  AAAGCGGGTG  600
601   CTGGAGTTGT  CACTAGAGGA  AGCTCGACAA  CTTGGGCATA  ACTACATTGG  TTCAGAGCAC  660
661   CTTTTGCTTG  GTCTACTTCG  TGAAGGGGAG  GGTGTGGCAG  CTCGTGTCTT  GGAGAATTTG  720
721   GGTGCAGATC  CTAGTAATAT  ACGGACACAG  GTTATACGTA  TGGTCGGGGA  AAACAATGAA  780
781   GTCACAGCAA  GCGTTGGTGG  GGGAAGCAGC  GGAAACAGCA  AAATGCCAAC  ACTTGAAGAG  840
841   TATGGGACTA  ACTTAACTAA  ACTAGCAGAG  GAGGGTAAAC  TGGATCCGGT  TGTTGGAAGG  900
901   CAGCCACAGA  TCGAACGAGT  GGTCCAGATC  TTGGCTCGAA  GAACCAAGAA  CAACCCATGT  960
961   CTTATTGGAG  AACCTGGAGT  TGGTAAGACG  GCAATAGCAG  AAGGACTTGC  ACAGCGAATA  1020
1021  GCTAGTGGTG  ATGTTCCTGA  AACAATCGAG  GGGAAGACGG  TTATAACCCT  TGATATGGGT  1080
1081  CTTCTAGTGG  CTGGAACGAA  ATACCGTGGA  GAGTTCGAGG  AAAGATTGAA  GAAGCTTATG  1140
1141  GAGGAAATCA  GGCAAAGTGA  TGAGATAATT  CTGTTTATTG  ATGAAGTGCA  CACGCTCATC  1200
1201  GGTGCAGGAG  CCGCTGAAGG  TGCGATCGAT  GCTGCTAACA  TCTTAAAGCC  AGCTCTAGCA  1260
1261  AGAGGTGAAT  TGCAGTGTAT  TGGTGCAACA  ACAATTGATG  AGTACAGGAA  ACACATTGAG  1320
1321  AAAGATCCTG  CATTGGAGAG  ACGGTTCCAG  CCTGTGAAAG  TACCTGAACC  AACTGTAGAA  1380
1381  GAAGCTATAC  AGATTTTGCA  AGGTCTGCGT  GAGCGCTATG  AGATCCACCA  CAAACTTCGA  1440
1441  TACACTGATG  AAGCCTTGGT  TGCTGCTGCA  CAATTGTCAC  ATCAGTACAT  CAGTGATCGG  1500
1501  TTTCTTCCCG  ACAAAGCGAT  TGACTTGATT  GATGAAGCTG  GGTCTCGGGT  TCGACTACGC  1560
1561  CATGCTCAGC  TTCCTGAGGA  AGCTAGAGAG  CTTGAAAAGC  AACTCAGGCA  AATCACCAAA  1620
1621  GAGAAGAATG  AAGCTGTGCG  AAGCCAAGAC  TTCGAGATGG  CTGGTTCTCA  TCGTGACCGT  1680
1681  GAAATAGAGC  TCAAGGCTGA  GATAGCTAAT  GTTTTATCTC  GAGGCAAAGA  AGTGGCCAAA  1740
1741  GCCGAGAATG  AAGCTGAGGA  AGGAGGACCT  ACTGTCACAG  AATCTGACAT  CCAACACATC  1800
1801  GTCGCCACCT  GGACAGGAAT  CCCGGTAGAG  AAAGTCTCGT  CTGATGAATC  TAGCCGTCTT  1860
1861  CTCCAAATGG  AGCAGACCCT  TCACACAAGA  GTCATTGGCC  AAGATGAAGC  CGTTAAAGCA  1920
1921  ATCAGTCGGG  CTATCCGCCG  TGCCCGTGTT  GGGCTCAAAA  ACCCAAACCG  TCCTATCGCC  1980
1981  AGTTTCATCT  TCTCTGGTCC  AACTGGTGTT  GGGAAATCAG  AGCTTGCTAA  GGCCTTGGCT  2040
2041  GCTTACTACT  TCGGTTCAGA  AGAAGCAATG  ATCCGTCTTG  ACATGAGTGA  GTTCATGGAA  2100
2101  CGACACACTG  TTTCGAAACT  CATCGGTTCA  CCTCCTGGTT  ACGTAGGATA  CACAGAAGGA  2160
2161  GGTCAGTTAA  CAGAGGCGGT  TCGACGCAGG  CCCTACACTC  TTGTTCTCTT  TGACGAAATT  2220
2221  GAGAAAGCAC  ATCCCGATGT  TTTCAACATG  ATGCTTCAGA  TCCTAGAAGA  CGGTAGACTA  2280
2281  ACTGATAGCA  AAGGAAGAAC  TGTCGATTTC  AAGAACACGC  TTCTGATCAT  GACTTCAAAC  2340
2341  GTAGGGAGCA  GCGTGATCGA  AAAAGGTGGT  AGAAGAATTG  GGTTTGATCT  TGACCACGAC  2400
2401  GAGAAAGACA  GCAGTTACAA  CAGAATCAAG  AGCTTAGTGA  CTGAGGAACT  AAAACAGTAT  2460
2461  TTCAGACCAG  AGTTCTTGAA  CAGGTTAGAT  GAGATGATTG  TTTTCAGACA  GTTAACAAAG  2520
2521  CTGGAAGTCA  AGGAGATTGC  TGATATAATG  CTTAAAGAAG  TGGTGGCGAG  ACTTGAGGTC  2580
2581  AAAGAGATTG  AGCTTCAGGT  GACTGAGAGG  TTTAAAGAGA  GAGTGGTGGA  TGAAGGATTC  2640
2641  GACCCGAGTT  ATGGTGCGAG  GCCACTTAGA  CGTGCAATAA  TGAGGCTTTT  GGAGGATAGT  2700
2701  ATGGCGGAGA  AGATGCTTTC  AAGGGACATT  AAAGAAGGAG  ATTCTGTGAT  TGTTGATGTT  2760
2761  GATGCCGAAG  GAAGTGTGGT  TGTGTTGAGT  GGTACCACTG  GACGTGTTGG  TGGTTTTGCT  2820
2821  GCTGAAGAAG  CCATGGAAGA  TCCTATTCCA  ATATTGTAG  2859

▼ KEYWORD


ID
Family
Alternative splicing
ATP-binding
Chaperone
Chloroplast
Complete proteome
Hydrolase
Membrane
Nucleotide-binding
Plastid
Reference proteome
Repeat
Transit peptide

▼ GENE ONTOLOGY


ID
Classification
Description
Cellular Component
Chloroplast
Cellular Component
Chloroplast envelope
Cellular Component
Chloroplast membrane
Cellular Component
Chloroplast stroma
Cellular Component
Membrane
Cellular Component
Mitochondrion
Cellular Component
Plastid
Cellular Component
Plastid stroma
Molecular Function
ATP binding
Molecular Function
ATPase activity
Molecular Function
Protein homodimerization activity
Biological Process
Chloroplast organization
Biological Process
Protein hexamerization
Biological Process
Protein import into chloroplast stroma
Biological Process
Protein metabolic process

▼ KEGG



▼ ANNOTATION


Disorder
IUPred2A
Physicochemical
AAindex
Function
CGDB
Localization
COMPARTMENTS

▼ ORTHOLOGY


DrLLPS IDOrganismIdentityE-valueScore
LLPS-Arl-1056Arabidopsis lyrata95.70.01679
LLPS-Bro-2733Brassica oleracea88.90.01540
LLPS-Brn-1020Brassica napus88.80.01536
LLPS-Brr-2460Brassica rapa88.780.01540
LLPS-Thc-1774Theobroma cacao87.930.01458
LLPS-Sot-2198Solanum tuberosum86.960.01419
LLPS-Glm-3010Glycine max86.620.01446
LLPS-Phv-1974Phaseolus vulgaris86.610.01441
LLPS-Nia-1812Nicotiana attenuata86.50.01408
LLPS-Ors-2283Oryza sativa86.220.0 681
LLPS-Sol-1622Solanum lycopersicum85.780.0 706
LLPS-Sob-1887Sorghum bicolor84.980.01380
LLPS-Dac-0668Daucus carota84.680.01431
LLPS-Brd-2195Brachypodium distachyon84.470.01378
LLPS-Orgl-1203Oryza glumaepatula84.390.01380
LLPS-Php-1803Physcomitrella patens84.130.01371
LLPS-Coc-0422Corchorus capsularis83.750.0 731
LLPS-Hov-1241Hordeum vulgare83.540.01378
LLPS-Orm-0862Oryza meridionalis83.390.01352
LLPS-Sei-1217Setaria italica83.390.01384
LLPS-Tra-0258Triticum aestivum83.260.01377
LLPS-Orbr-2322Oryza brachyantha83.150.01386
LLPS-Orr-2078Oryza rufipogon83.040.01383
LLPS-Org-2230Oryza glaberrima83.040.01383
LLPS-Orb-2026Oryza barthii83.040.01383
LLPS-Orni-2263Oryza nivara83.040.01383
LLPS-Chc-0431Chondrus crispus74.820.01143
LLPS-Cym-0728Cyanidioschyzon merolae71.150.01050
LLPS-Gas-1035Galdieria sulphuraria70.620.01100
LLPS-Vir-1159Vigna radiata66.139e-95 318
LLPS-Yal-0808Yarrowia lipolytica65.55e-94 321
LLPS-Nec-0846Neurospora crassa56.034e-75 271
LLPS-Scc-0924Schizosaccharomyces cryophilus55.113e-81 288
LLPS-Map-0065Magnaporthe poae55.072e-74 269
LLPS-Mao-0995Magnaporthe oryzae55.073e-79 283
LLPS-Gag-1207Gaeumannomyces graminis54.457e-79 281
LLPS-Ten-3401Tetraodon nigroviridis54.441e-27 114
LLPS-Blg-0261Blumeria graminis54.353e-76 274
LLPS-Tru-1445Triticum urartu54.343e-110 366
LLPS-Trv-0833Trichoderma virens54.327e-71 258
LLPS-Chr-0207Chlamydomonas reinhardtii54.246e-111 374
LLPS-Asfu-0935Aspergillus fumigatus54.214e-77 277
LLPS-Nef-0618Neosartorya fischeri54.212e-77 278
LLPS-Scs-0524Sclerotinia sclerotiorum53.191e-77 278
LLPS-Ast-1151Aspergillus terreus53.168e-82 290
LLPS-Pot-1726Populus trichocarpa52.89e-88 306
LLPS-Asc-0102Aspergillus clavatus52.793e-77 277
LLPS-Cogr-0056Colletotrichum graminicola52.773e-72 262
LLPS-Coo-0375Colletotrichum orbiculare52.682e-77 277
LLPS-Fuo-0585Fusarium oxysporum52.612e-72 263
LLPS-Aso-0808Aspergillus oryzae52.468e-82 290
LLPS-Asf-0658Aspergillus flavus52.468e-82 290
LLPS-Via-1096Vigna angularis52.135e-87 305
LLPS-Asni-0338Aspergillus niger52.134e-81 288
LLPS-Lem-0864Leptosphaeria maculans51.859e-79 281
LLPS-Mel-1210Melampsora laricipopulina51.81e-71 260
LLPS-Gor-1005Gossypium raimondii51.521e-87 306
LLPS-Fuv-0143Fusarium verticillioides51.241e-72 263
LLPS-Cus-1314Cucumis sativus50.980.0 672
LLPS-Ori-0392Oryza indica50.743e-85 299
LLPS-Met-1362Medicago truncatula50.560.0 668
LLPS-Prp-2273Prunus persica50.490.0 668
LLPS-Fus-0716Fusarium solani50.317e-72 261
LLPS-Abg-1806Absidia glauca49.42e-43 176
LLPS-Osl-1429Ostreococcus lucimarinus49.252e-107 358
LLPS-Miv-0078Microbotryum violaceum49.225e-79 282
LLPS-Put-1003Puccinia triticina49.116e-37 154
LLPS-Sem-2027Selaginella moellendorffii48.80.0 654
LLPS-Cog-0117Colletotrichum gloeosporioides48.668e-66 243
LLPS-Sac-0692Saccharomyces cerevisiae48.633e-73 265
LLPS-Tum-0582Tuber melanosporum48.560.0 591
LLPS-Asg-1467Ashbya gossypii48.370.0 606
LLPS-Pug-0177Puccinia graminis47.176e-71 258
LLPS-Kop-1085Komagataella pastoris47.084e-80 285
LLPS-Viv-1634Vitis vinifera45.980.0 647
LLPS-Orl-0943Oryzias latipes45.411e-46 179
LLPS-Amt-1529Amborella trichopoda45.30.0 613
LLPS-Lep-0422Leersia perrieri45.040.0 560
LLPS-Hea-0529Helianthus annuus44.670.0 656
LLPS-Orp-1021Oryza punctata44.110.0 571
LLPS-Zem-0889Zea mays43.840.0 559
LLPS-Scj-0156Schizosaccharomyces japonicus43.720.0 555
LLPS-Phn-0963Phaeosphaeria nodorum42.962e-172 533
LLPS-Dar-0309Danio rerio42.563e-50 184
LLPS-Trr-0808Trichoderma reesei42.538e-172 532
LLPS-Mae-0209Manihot esculenta42.522e-168 522
LLPS-Pyt-0513Pyrenophora teres42.521e-168 523
LLPS-Pytr-0785Pyrenophora triticirepentis42.399e-168 521
LLPS-Beb-0114Beauveria bassiana42.291e-170 529
LLPS-Usm-1081Ustilago maydis42.082e-173 536
LLPS-Icp-3117Ictalurus punctatus42.082e-45 172
LLPS-Scf-2418Scleropages formosus41.983e-50 186
LLPS-Scp-0309Schizosaccharomyces pombe41.963e-180 553
LLPS-Ved-0605Verticillium dahliae41.851e-167 521
LLPS-Spr-0561Sporisorium reilianum41.425e-175 540
LLPS-Dos-0315Dothistroma septosporum40.91e-171 532
LLPS-Mua-0196Musa acuminata40.65e-157 491
LLPS-Crn-0669Cryptococcus neoformans38.969e-148 468
LLPS-Asn-0154Aspergillus nidulans38.915e-134 432
LLPS-Pes-2325Pelodiscus sinensis38.753e-53 199
LLPS-Fia-0060Ficedula albicollis38.597e-54 202
LLPS-Caj-3549Callithrix jacchus38.592e-53 203
LLPS-Poa-0622Pongo abelii38.262e-53 202
LLPS-Mam-2023Macaca mulatta38.262e-53 202
LLPS-Cas-3171Carlito syrichta38.262e-53 202
LLPS-Nol-2938Nomascus leucogenys38.267e-54 202
LLPS-Hos-3083Homo sapiens38.263e-53 202
LLPS-Pat-0742Pan troglodytes38.263e-53 202
LLPS-Otg-4006Otolemur garnettii38.263e-53 202
LLPS-Rhb-0218Rhinopithecus bieti38.263e-53 202
LLPS-Bot-2096Bos taurus38.261e-53 203
LLPS-Ova-4364Ovis aries38.264e-53 203
LLPS-Mal-4455Mandrillus leucophaeus38.262e-53 203
LLPS-Chs-3598Chlorocebus sabaeus38.262e-53 202
LLPS-Cea-4275Cercocebus atys38.262e-53 202
LLPS-Gog-1785Gorilla gorilla38.264e-53 202
LLPS-Lac-3981Latimeria chalumnae38.161e-53 197
LLPS-Mea-1371Mesocricetus auratus38.11e-40 158
LLPS-Dio-3829Dipodomys ordii38.039e-41 158
LLPS-Ora-3114Ornithorhynchus anatinus38.032e-41 160
LLPS-Cap-2885Cavia porcellus37.926e-53 201
LLPS-Eqc-3803Equus caballus37.922e-53 203
LLPS-Ict-1055Ictidomys tridecemlineatus37.926e-53 201
LLPS-Urm-1617Ursus maritimus37.922e-53 202
LLPS-Loa-1378Loxodonta africana37.926e-54 204
LLPS-Ran-3757Rattus norvegicus37.925e-53 201
LLPS-Mup-4369Mustela putorius furo37.923e-53 202
LLPS-Aim-0168Ailuropoda melanoleuca37.922e-53 203
LLPS-Mum-1442Mus musculus37.925e-53 201
LLPS-Sah-3272Sarcophilus harrisii37.922e-53 199
LLPS-Fud-0247Fukomys damarensis37.927e-53 201
LLPS-Tag-1382Taeniopygia guttata37.878e-53 195
LLPS-Caf-1762Canis familiaris37.582e-53 203
LLPS-Fec-4339Felis catus37.582e-53 203
LLPS-Orc-3421Oryctolagus cuniculus37.589e-53 201
LLPS-Myl-0372Myotis lucifugus37.581e-53 204
LLPS-Anp-1789Anas platyrhynchos37.557e-38 150
LLPS-Anc-0653Anolis carolinensis37.52e-48 186
LLPS-Mod-4047Monodelphis domestica37.224e-52 199
LLPS-Sus-3217Sus scrofa37.223e-54 205
LLPS-Gaa-1222Gasterosteus aculeatus37.123e-49 190
LLPS-Scm-3500Scophthalmus maximus35.915e-50 192
LLPS-Leo-2831Lepisosteus oculatus35.829e-51 193
LLPS-Pap-2393Pan paniscus35.696e-45 177
LLPS-Man-3804Macaca nemestrina35.696e-45 177
LLPS-Paa-2339Papio anubis35.698e-45 177
LLPS-Maf-3768Macaca fascicularis35.695e-45 177
LLPS-Orn-4126Oreochromis niloticus34.759e-52 196
LLPS-Asm-0920Astyanax mexicanus34.291e-52 199
LLPS-Aon-3173Aotus nancymaae34.015e-42 168
LLPS-Tar-1942Takifugu rubripes33.934e-49 189
LLPS-Pof-4133Poecilia formosa33.92e-50 193
LLPS-Xim-3177Xiphophorus maculatus33.622e-49 191
LLPS-Xet-3261Xenopus tropicalis32.695e-37 153